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??????????????????龥?????

????????????????Helianthus tuberosus L.

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???????????? ?????????????1-3m???????????????????????????????????????????????????????????????????????????????????????????????Σ???10-15cm????3-9cm????????????????Ш?Σ?????о?????????????汻???????3???????????????????????????5-9cm????1-2??????????ε??????????????λ?????????Σ??????????????????????????????????????????????????????????????????????5?????Ш?Σ??????????2-4???????????????8-10???

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??????????????????????????????,?????????????????????,????????,????????????????,??10-15cm,??3-9cm,3??,???????,?±????????,??????,????????,??????Ш??,????????????

??????????????????(inulin),????1F-??-D-????????(sucrose 1F-??-D-fructosyltransferase),?????-1,5-??????????(ribulose-l,5-bisphosphatecarboxylase),????????(polyphenoloxidase),???????(inulase),????????(fructo-oligosaccharides)?? ??????????(heliangine),?????????(tagitinin)E,???????????(erioflorin)??????????????????(leptocarpin),14-???????????(14-hydroxyleptocarpin),???????(budlein)A,???????A???????(budlein A tighate),???????A2-?????????(budlein A2-methylbutyrate),??????? A??????(budleinA isobutyrate),???????A??????????(budlein A methylacry-late),4,5-??-???????A??????(4,5-iso-budlein Aisobutyrate),4,15-??-??????????????????????(4,15-iso-a-tripliciolide isobutyrate),4,15-??????????????????????????(4,15-iso-atripliciolide methyl acrylate,4,15-???????????????????????(4,15-iso-atripliciolide tiglate),4,15-????????????????????????(4,15-iso-atripliciolide angelate),3-???-???????????????????????(3-hydroxy-atripliciolide tiglate),8??-14?????????????(8??,14-dihydroxy costunolide),????????????????(desacetyleupaserrin),14-??????????????黷??????????(14-acetoxy tlthifolin epoxyangelate), 1??, 2-??????????????(l??,2-dihydroxy pinnatifidin),?????????(pyrenocin)A??B,???????(jasmonic acid),?????-D-?????????龥?????(Me??-D-glucopyranosyl tuberonate),?????-D-??????????????????(Me??-D-glucopyranosyl helianthenate)A??B??C??D??E??F,????龥??????????(Me tuberonic acid giucoside)?? ????????????????????(helianthol)A??????????????????Ц?-?????(??-bisabolene)??